Officials from Saturday's USC-Virginia game in Los Angeles were reprimanded and removed from a future assignment by the Pac-10 offices Monday.
At issue was an illegal block call made on a Virginia fake punt in the second quarter.
The play would have given the Cavaliers the ball deep in USC territory, but the penalty nullified it.
USC won 17-14.
A team is indeed allowed to block below the waist on a fake punt.
Officials apologized to Virginia on Saturday.
"We know mistakes are made in games," Pac-10 commissioner Larry Scott said. "But we will not tolerate our officials misinterpreting the rules."
Exam: Penn player had brain disease
PHILADELPHIA - A detailed examination of the brain of a Penn football player who committed suicide showed he had the same disease caused by hard hits that has been associated with NFL players.
The New York Times first reported Monday that Owen Thomas, 21, was in the early stages of chronic traumatic encephalopathy, or CTE, a disease that has been linked to depression and impulse control, primarily among NFL players.
Thomas, a former Penn captain, was found dead in April at his off-campus apartment. He had apparently hanged himself.
The research on Thomas' brain tissue was done at Boston University, which has done much of the work on the impact of big hits on football players that has gained attention in recent years. Dr. Robert Stern, a director of the Boston University group, confirmed the findings to The Associated Press but otherwise declined comment.
• Alabama tailback Mark Ingram practiced Monday, wearing a brace on his left knee.
The Heisman Trophy winner appears on track to return Saturday against Duke after missing the first two games with a knee injury.
• The Heisman Trophy Trust is expected to hold its monthly meeting today, but there has been no indication from the group about whether a decision on Reggie Bush and the 2005 award is coming soon.
Heisman spokesman Tim Henning said Monday the eight-member trust had no plans to make any announcements at this time.
• Missouri linebacker Will Ebner and long snapper Beau Brinkley have been reinstated from two-game disciplinary suspensions for alcohol-related arrests.